Against the wave of accelerating global transformation, a new political force known as the tech-right has rapidly emerged on the U.S. social and political stage. Driven by the advancement of economic globalization, digitalization, and AI transformation, this force is grounded in accelerationism as its philosophical foundation, guided by right-wing conservatism as its dominant ideology, and adheres to anti-establishment and anti-system political logic. It is now spearheading a politically transformative shift in the U.S. of historic significance. In essence, this transformation represents a political reckoning with progressive America since the era of Roosevelt. By engaging in U.S. party politics, electoral politics, and governance, it has ushered in a new era dominated by right-wing conservatism. To grasp the political potential of the tech-right, one must examine its social foundation and ecological reason from the perspective of the global transformation ecosystem in which American politics operates. From the standpoint of ecological institutionalism, the historical significance of the tech-right’s rise lies in its response to the essential changes in U.S. modes of production during the digital and AI era. These changes will lead to substantial transformations in the U.S. constitutional structure over an extended period, driving the nation from a progressive era dominated by liberal democratic consensus into a conservative era led by an “America First” consensus. It thus pioneers a new future for American politics.